-Growing conditions are not so optimal, so the tree canopy is lower (10-30m) than in the tropical rain forest and the trees less dense where drought is more extreme.
-The undergrowth is dense and tangled because of greater light penetration.
-Lianas (woody vines) are much less common than in the rain forest, not such an important growth form where light is less limiting and also perhaps highly susceptible to desiccation.
-Drought-resistant epiphytes (orchids, bromeliads and cacti) may be abundant.
-The trees have thicker, more ridged, bark; deeper roots without buttresses; much more variable leaves, including many compound-leaved legumes; and more species with thorns.